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| Matthew Broderick | Despereaux |
| Dustin Hoffman | Roscuro |
| Emma Watson | Pea |
| Tracey Ullman | Mig |
| Kevin Kline | Andre |
| William H. Macy | Lester |
| Stanley Tucci | Boldo |
| Ciarán Hinds | Botticelli |
| Robbie Coltrane | Gregory |
| Frances Conroy | Antoinette |
| Tony Hale | Furlough |
| Frank Langella | Mayor |
| Richard Jenkins | Principal |
| Christopher Lloyd | Hovis |
Once a thriving place of sunshine and delicious soup, the faraway kingdom of Dor has fallen into sadness. An accident leaves the king brokenhearted; the sunshine fades, and the soup disappears Into this world is born Despereaux, a book-loving mouse and friend to Princess Pea . When Pea is kidnapped, Despereaux must find a knight's brave heart within his tiny body and rescue his friend.
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