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| Martin Freeman | Arthur Dent |
| Mos Def | Ford Prefect |
| Sam Rockwell | Zaphod Beeblebrox |
| Zooey Deschanel | Tricia McMillan ("Trillian") |
| Warwick Davis | Marvin the Paranoid Android |
| Alan Rickman | Marvin the Paranoid Android |
| John Malkovich | Humma Kavula |
| Bill Nighy | Slartibartfast |
| Thomas Lennon | Eddie the Computer |
| Helen Mirren | Deep Thought |
| Anna Chancellor | Questular Rontok |
| Stephen Fry | The Book |
| Bill Bailey | Sperm Whale |
| Mason Ball | Vogon Minion |
| William Todd Jones | Vogon Captain |
| Barry Dowden | Stunt Builder |
| Simon Jones | Magrathean Announcer |
A human and his extraterrestrial friend begin an interstellar journey after the destruction of Earth.
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