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| William Shatner | Captain James Tiberius Kirk |
| Leonard Nimoy | Captain Spock |
| DeForest Kelley | Commander Leonard H. McCoy, M.D. |
| James Doohan | Captain Montgomery 'Scotty' Scott |
| Walter Koenig | Commander Pavel Andreievich Chekov |
| Nichelle Nichols | Commander Nyota Uhura |
| George Takei | Captain Hikaru Sulu |
| Kim Cattrall | Lieutenant Valeris |
| David Warner | Chancellor Gorkon |
| Christopher Plummer | General Chang |
| Mark Lenard | Vulcan Ambassador Sarek |
| Grace Lee Whitney | Commander Janice Rand |
| Brock Peters | Admiral Cartwright |
| Leon Russom | Chief in Command |
| Kurtwood Smith | Federation President |
| Rosanna DeSoto | Chancellor Azetbur |
| John Schuck | Klingon Ambassador |
| Michael Dorn | Colonel Worf |
| Paul Rossilli | Brigadier Kerla |
| Christian Slater | Excelsior Communications Officer |
Capt. James Kirk and the crew of the USS Enterprise are carrying Klingon Chancellor Gorkon to Earth to negotiate a peace treaty with the United Federation of Planets. The ship appears to fire on a Klingon vessel, and Gorkon is killed in the subsequent confusion. Kirk and the ship's doctor, Leonard McCoy, are arrested for murder, leaving Spock to figure out who is behind the attack and save the negotiations.
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