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| Bruce Willis | RJ |
| Garry Shandling | Verne |
| Steve Carell | Sammy the Squirrel |
| William Shatner | Ozzie the Possum |
| Allison Janney | Gladys |
| Wanda Sykes | Stella the Skunk |
| Thomas Haden Church | Dwayne LaFontaine |
| Avril Lavigne | Heather |
| Eugene Levy | Lew the Porcupine |
| Catherine O'Hara | Penny the Porcupine |
| Nick Nolte | Vincent |
When Verne and fellow woodland friends awake from winter's hibernation, they find they have some new neighbors: humans, and RJ, an opportunistic raccoon who shows the others how to exploit the suburban bounty laid out before them.
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