|Jada Pinkett Smith||Gloria|
|Sacha Baron Cohen||King Julien XIII|
|Frances McDormand||Capitaine Chantel DuBois|
|Cedric the Entertainer||Maurice|
|Conrad Vernon||Mason the Chimp|
Animal pals Alex, Marty, Melman and Gloria are still trying to make it back to New York's Central Park Zoo. They are forced to take a detour to Europe to find the penguins and chimps who broke the bank at a Monte Carlo casino. When French animal-control officer Capitaine Chantel DuBois picks up their scent, Alex and company are forced to hide out in a traveling circus.