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| Peter O'Toole | The Duke of Rudling |
| Samantha Morton | Sarah Carraclough |
| John Lynch | Sam Carraclough |
| Peter Dinklage | Rowlie |
| Steve Pemberton | Eddie Hynes |
| Jemma Redgrave | Daisy |
| Jonathan Mason | Joe Carraclough |
| Hester Odgers | Cilla |
| Edward Fox | Col. Hutton |
| John Standing | Col. French |
| Gregor Fisher | Mapes |
| Kelly Macdonald | Jeanie |
| Jamie Lee | Tom |
| Robert Hardy | Mr. Justice Murray |
| Nicholas Lyndhurst | Buckles |
Young Joe Carraclough and his family love their faithful collie, Lassie. However, when Joe's father loses his job in the mine, he must sell the dog to the Duke of Rudling, who covets the collie for his granddaughter . After the duke moves to northern Scotland to avoid the threat of World War II, Lassie escapes and undertakes a 500-mile journey to reunite with her family.
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