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| Christopher Lloyd | Grandpa |
| Kelly Ripa | Nat's mom |
| Nicollette Sheridan | Nadia |
| Tim Curry | Yegor |
| Trevor Gagnon | Nat |
| Philip Daniel Bolden | IQ |
| David Gore | Scooter |
| Ed Begley Jr. | Poopchev |
| Robert Patrick | Louie |
| Adrienne Barbeau | Scooter's mom |
| Buzz Aldrin | Himself |
| Sandy Simpson | Commander Armstrong |
| Eddie Frierson | Commander Aldrin |
| David Cowgill | Commander Collins |
| Steve Kramer | Leonid |
| Mimi Maynard | IQ's mom |
Nat the fly loves listening to his grandfather tell stories of adventures from his youth. Desperate to experience some derring-do of his own, Nat convinces two of his friends to stow away on Apollo 11. The high-flying trio have the adventure of a lifetime when they must prevent a Russian bug from sabotaging the mission.
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