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| Ben Stiller | Mel Coplin |
| Patricia Arquette | Nancy Coplin |
| Tea Leoni | Tina Kalb |
| Alan Alda | Richard Schlichting |
| Mary Tyler Moore | Mrs. Coplin |
| George Segal | Mr. Coplin |
| Lily Tomlin | Mary Schlichting |
| Josh Brolin | Tony |
| Richard Jenkins | Paul |
| Celia Weston | Valerie Swaney |
| Glenn Fitzgerald | Lonnie Schlichting |
| Beth Ostrosky | Jane |
| Cynthia Lamontagne | Sandra |
| David Patrick Kelly | Fritz Boudreau |
| John Ford Noonan | Mitch |
| Charlet Oberly | B&B Lady |
Adopted as a child, new father Mel Colpin decides he cannot name his son until he knows his birth parents, and determines to make a cross-country quest to find them. Accompanied by his wife, Nancy, and an inept yet gorgeous adoption agent, Tina, he departs on an epic road trip that quickly devolves into a farce of mistaken identities, wrong turns, and overzealous and love-struck ATF agents .
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