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| George Clooney | Mr. Fox |
| Meryl Streep | Mrs. Fox |
| Jason Schwartzman | Ash |
| Bill Murray | Badger |
| Wally Wolodarsky | Kylie |
| Eric Anderson | Kristofferson |
| Michael Gambon | Franklin Bean |
| Willem Dafoe | Rat |
| Owen Wilson | Coach Skip |
| Jarvis Cocker | Petey |
| Wes Anderson | Weasel |
| Karen Duffy | Linda Otter |
| Robin Hurlstone | Walter Boggis |
| Hugo Guinness | Nathan Bunce |
| Helen McCrory | Mrs. Bean |
| Roman Coppola | Squirrel Contractor |
| Juman Malouf | Agnes |
| Jeremy Dawson | Beaver's Son |
| Garth Jennings | Bean's Son |
After 12 years of bucolic bliss, Mr. Fox breaks a promise to his wife and raids the farms of their human neighbors, Boggis, Bunce and Bean. Giving in to his animal instincts endangers not only his marriage but also the lives of his family and their animal friends. When the farmers force Mr. Fox and company deep underground, he has to resort to his natural craftiness to rise above the opposition.
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