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| Owen Wilson | Lightning McQueen |
| Larry the Cable Guy | Mater |
| Michael Caine | Finn McMissile |
| Emily Mortimer | Holley Shiftwell |
| John Turturro | Francesco Bernoulli |
| Eddie Izzard | Miles Axlerod |
| Bonnie Hunt | Sally |
| Tony Shalhoub | Luigi |
| Cheech Marin | Ramone |
| Jason Isaacs | Siddeley/Leland Turbo |
| Thomas Kretschmann | Professor Z |
| Peter Jacobson | Acer |
| Joe Mantegna | Grem |
| Franco Nero | Uncle Topolino |
| Vanessa Redgrave | Mama Topolino/The Queen |
| Bruce Campbell | Rod ``Torquie'' Redline |
| Michel Michelis | Tomber |
| Jeff Gordon | Jeff Gorvette |
| Darrell Waltrip | Darrell Cartrip |
| Lewis Hamilton | Lewis Hamilton |
| David Hobbs | David Hobbscap |
| Jenifer Lewis | Flo |
| Jeff Garlin | Otis |
| Sig Hansen | Crabby |
Racecar Lightning McQueen and his tow-truck buddy, Mater, have had their share of adventures together but nothing like what they experience when they head overseas for the first-ever World Grand Prix. While Lightning has his eye on the road to the championship, Mater takes a detour to international espionage when he meets British master spy Finn McMissile and stunning spy-in-training Holley Shiftwell .
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